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Terrestrial Gastropods of the Columbia Basin, British Columbia

Key to Terrestrial Gastropods of British Columbia
47a
Major axial sculpture of more or less regularly spaced grooves, with smooth, flattened, raised areas between (see figure below)  -  48
47b
Shell smoothish or with axial sculpture not as above  -  49


Cross-section (perpendicular to the axial grooves) of the shell wall of Nesovitrea
showing characteristic grooves and flattened areas between.

 
48a
Shell nearly colourless with a greenish tinge and smaller, 3.5-4.3 mm (3½-4½ whorls)  -  Nesovitrea binneyana (Figure)
48b
Shell amber-coloured and larger, 4.6-5.2 mm (3½-4 whorls)  -  Nesovitrea electrina (Figure)

49a
Shell with widely and more or less regularly spaced (at least on the whorls of the spire), sharply angular or lamellar axial ribs  -  50
49b
Shell smoothish or with axial sculpture closely and either regularly or irregularly spaced  -  55

50a
Shell white, grey or yellowish grey  -  51
50b
Shell yellowish brown, brown or reddish brown  -  53

51a
Lip flared but thin  -  52
51b
Lip at least slightly thickened  -  Vallonia gracilicosta

52a
Shell width to about 2.2 mm  -  Vallonia cf. perspectiva (Figure)
52b
Shell width 2.5-3.4 mm  -  Vallonia cyclophorella (Figure)

53a
Spire almost flat  -  Planogyra clappi
53b
Spire moderately raised  -  54

54a
Umbilicus about 20-25% of the width of the shell; protoconch initially smooth, then with rather widely spaced spiral striae  -  Paralaoma caputspinulae
54b
Umbilicus larger, greater than 30% of the width of the shell; protoconch smooth  -  36

55a
Lip thickened within by a callus  -  56
55b
Lip not thickened within  -  57

56a
Last whorl near the aperture distinctly increasing in width much more than previously; outline of the shell ovate; umbilicus elongate, elliptically spiral; outer lip gradually expanded, not at a right angle  -  Vallonia excentrica (Figure)
 
      Introduced to British Columbia (Forsyth in press)
56b
Last whorl near the aperture increasing in width only a little more than previously; outline of the shell almost circular; umbilicus regularly spiral; outer lip abruptly flared outward at a right angle Vallonia pulchella (Figure)
Introduced to British Columbia (Forsyth in press)

57a
Whorls rapidly enlarging; umbilicus minute  -  Vitrina pellucida (Figure)
57b
Whorls enlarging more slowly; umbilicus about 15-30% of the width of the shell  -  58

58a
Shell whitish or colourless  -  59
58b
Shell yellowish, yellowish brown, brown or reddish brown  -  63

59a
Umbilicus about 15% of the shell width  -  Vitrea contracta
Introduced to British Columbia (Forsyth in press)
59b
Umbilicus about 25-30% of the shell width  -  60

60a
Axial sculpture of fine, close riblets  -  Striatura pugetensis
60b
Axial sculpture more or less non-existent; fine incremental striae only  -  61

61a
Umbilicus about 30% of the shell width; spire moderately elevated  -  Hawaiia minuscula
61b
Umbilicus about 25% of the shell width; spire very low  -  62

62a
Width 4-5 mm; 5-5¼ whorls; SE British Columbia  -  Microphysula ingersolli (Figure)
62b
Width 3.6-4.4 mm; 4-4½ whorls; Coast and Cascade Mountains and Vancouver Island Range  -  Microphysula cookei

63a
Whorls rapidly enlarge in width: last whorl at least twice the width of the penultimate whorl one-half coil back from aperture  -  64
63b
Whorls less rapidly enlarge in width: last whorl one-half coil back from aperture last whorl less than twice the width of the penultimate whorl  -  65

64a
Axial sculpture of fine, close riblets; protoconch with minute, close spiral striae; shell width to 1.7 mm (2¾-3 whorls); width of umbilicus about 30% of the width of the shell  -  Striatura pugetensis
64b
Axial sculpture weak and shell almost smooth; protoconch smooth; shell width greater than 5 mm (5-6 whorls); width of umbilicus about 17% of the width of the shell  -  Oxychilus alliarius
Introduced to British Columbia (Forsyth in press)

65a
Shell width up to 1.4 mm or 1.8 mm (3½-4½ whorls)  -  P. californicum & Punctum randolphii
P. californicum is generally slightly larger than P. randolphii.
65b
Shell larger  -  66

66a
Aperture narrow and crescent-shaped  -  67
66b
Aperture broader, more rounded  -  68

67a
Umbilicus tiny, pit-like  -  Euconulus fulvus (Figure)
67b
Umbilicus larger, about 10% of the width of the shell  -  Pristiloma chersinella (Figure)

68a
Animal bluish grey on tentacles and on back, lighter on sides and tail; shell width 5-6 mm (4½ whorls); shell with faint and extremely fine, microscopic spiral striae; generally inhabiting drier areas than the next species  -  Zonitoides arboreus (Figure)
68b
Animal blackish; shell width 6-7 mm (4½ whorls); microscopic spiral striae absent; generally inhabiting marshy areas  -  Zonitoides nitidus (Figure)
The spiral striae present on the shell surface of Z. arboreus are exceptionally fine. They should be looked for, using reflective light and a minimum of 30× magnification, between the larger axial riblets; however, not all specimens show striae clearly, even at this magnification, and in some specimens striae are apparently absent. Pigmentation of the animal is the most reliable distinguishing feature and should be used whenever known. Zonitoides arboreus is characteristically more depressed and with a wider umbilicus than Z. nitidus.

 

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