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Terrestrial Gastropods of the Columbia Basin, British Columbia

Key to Terrestrial Gastropods of British Columbia
17a
Parietal denticle present  -  18
17b
Parietal denticle absent  -  20

18a
Periostracal hairs usually persistent in the adult; shell width to about 7 mm  -  Cryptomastix germana
18b
Periostracal hairs not usually persistent in the adult; shell width greater than 12 mm  -  19

19a
Shell width 12-26 mm; basal lip denticle or swelling (at its thickest point) located relatively closer to the columella than the periphery; southwest coastal British Columbia  -  Cryptomastix devia
19b
Shell width 12-19 mm; basal lip denticle or swelling (at its thickest point) located relatively closer to the periphery than to the columellar; in southeast British Columbia  -  Cryptomastix mullani (Figure)

20a
Umbilicus absent  -  21
20b
Umbilicus open, although often slitlike or just a tiny pit  -  26

21a
Shell width greater than 15 mm and usually with colour spiral bands, but sometimes without  -  22
21b
Shell width less than 5 mm and without colour bands  -  23

22a
Aperture height less than its width; unicolour or typically banded, with bands continuous, seldom interrupted by axial streaks  -   Cepaea nemoralis
22b
Aperture height about equal to its width; banded, with bands usually interrupted by light-coloured axial streaks  -   Helix aspersa

23a
Aperture broadly and deeply rounded; whorls about 3  -  Vitrina pellucida (Figure)
23b
Aperture narrowly crescent-shaped; whorls 3½ or more  -  24

24a
Whorls with distinct axial grooves between the periphery and the suture in addition to very fine axial striae  -  Pristiloma stearnsii
24b
Whorls with fine axial striae (and sometimes spiral striae) only  -  25

25a
Shell waxy white or colourless; height of shell less than or equal to ½ of its width  -  Pristiloma johnsoni
25b
Shell brownish; height of shell greater than ½ its width  -  Pristiloma lansingi

26a
Shell width greater than or equal to 6 mm  -  27
26b
Shell width less than 6 mm  -  47

27a
Shell with colour bands  -  28
27b
Shell without colour bands  -  31

28a
Columellar lip expanded to almost close the umbilicus, leaving only a narrow slitlike opening  -  22
28b
Columellar lip overhanging the umbilicus, but not enough to leave only a narrow slitlike opening  -  29

29a
Shell brown with a narrow, pale, somewhat yellowish band at the periphery, a narrow, dark brown band above this and a dark brown base; shell rarely yellow with pale banding; animal with a reddish tinge; coastal British Columbia  -  Monadenia fidelis
29b
Shell greyish to brown, and banding not as above; animal not with reddish tinge; SE British Columbia  -  30

30a
Shell dark brown, generally with a lighter peripheral band obscurely bordered by a darker band on either side; this light band visible within the aperture  -  Anguispira kochi (Figure)
30b
Shell generally greyish or brownish grey with two or more (often interrupted) dark bands (but banding is highly variable and may be absent) and speckling; dark bands often visible within the aperture  -  Oreohelix strigosa & O. subrudis
O. subrudis usually has a more elevated spire than O. strigosa, but identification by this shell character can be unreliable.

31a
Periostracum with minute hairs  -  32
31b
Periostracum without hairs  -   33

32a
Lip flared, sometimes a little recurved, thin-edged  -  Vespericola columbianus
32b
Lip flared, strongly recurved, edge appearing thick due to the tight scroll-like folding back of the edge  -   Cryptomastix mullani (Figure)


Cross-section of the lip of Vespericola columbianus (top)
and Cryptomastix mullani (bottom)
33a
Shell with prominent, sharply defined regular axial ribs (at least on the upper surface of the whorls of the spire)  -  34
33b
Shell without axial ribs, or axial ribs weak or not regular  -  37

34a
Outer lip straightened or concave at shoulder in apertural view  -  35
34b
Outer lip more or less convex at shoulder in apertural view  -  36

35a
Beaded sculpture present on all whorls of the shell; width to 18 mm  -  Ancotrema sportella
35b
Beaded sculpture obsolete on the penultimate and/or last whorl; width to 27 mm  -  Ancotrema hybridum

36a
Axial ribs about as strong on the base as on the apical surface of the shell  -  Discus whitneyi
36b
Axial ribs absent or significantly weaker on the base than on the apical surface of the shell  -  Discus shimekii

37a
Lip strongly expanded and curved back to form a more or less broad, flattened face throughout its length  -  38
37b
Lip thin or slightly thickened but not greatly expanded into a flat face; the lip of Cepaea nemoralis is somewhat expanded at the periphery but not elsewhere along its length  -  40

38a
Sculpture of irregular, rounded axial ribs, strongest near the suture and usually disappearing below the periphery; width greater than 19 mm  -  39
38b
Sculpture of fine axial striae only; width 9-17 mm  -  40

39a
Shell width 21-31 mm; surface sculpture often malleated; apparently restricted in British Columbia to the vicinity of the Chilliwack Valley  -  Allogona townsendiana
39b
Shell width 19-24 mm; sculpture seldom malleated; southeast British Columbia  -  Allogona ptychophora

40a
Whorls about 3; shell pale, greenish, yellowish-green or nearly colourless  -  Vitrina pellucida
40b
Whorls about 4-6; shell amber-coloured or pale brown, often paler around the umbilicus  -  41

41a
Columellar lip expanded over to almost close the umbilicus, leaving only a narrow slitlike opening  -  Cepaea nemoralis
41b
Columellar lip may overhang the umbilicus, but not enough to leave only a narrow slitlike opening  -  42

42a
Axial sculpture rather rough and irregular and/or shell width usually greater than 16 mm  -  43
42b
Shell smoothish; width less than or equal to 16 mm  -  44

43a
Shell yellowish, olive green or darker; outer lip straightened at shoulder in apertural view  -  Haplotrema vancouverense
43b
Shell greyish white or occasionally more brownish; outer lip convex at shoulder  -  Oreohelix strigosa & O. subrudis
O. subrudis usually has a more elevated spire than O. strigosa, but identification by this shell character can be unreliable.

44a
Umbilicus about 25% of the shell width; shell with very fine, faint spiral striae  -  Aegopinella nitidula
Introduced to British Columbia (Forsyth et al. in prep.)
44b
Umbilicus about 15% of the shell width; shell without spiral striae  -  45

45a
Animal with a strong garlic odour when disturbed; shell to about 7 mm  -  Oxychilus alliarius
Introduced to British Columbia (Forsyth in press)
45b
Animal greyish, bluish-grey or nearly black, never with a garlic odour; shell greater than or equal to 7 mm  -  46

46a
Shell to about 10 mm in width; last whorl regularly expanding; shell usually very glossy and with only very weak striae if any; animal pale grey with minute brown specks on the edge of the mantle (evident near the pneumostome on the right side, and on the left side through the shell)  -  Oxychilus cellarius
Introduced to British Columbia (Forsyth in press)
46b
Shell larger, to 12-16 mm in width; last whorl more rapidly expanded near the aperture so that it appears more than twice as wide as the previous whorl; shell usually less glossy and more striate than above species; animal dark bluish grey  -  Oxychilus draparnaudi
Introduced to British Columbia (Forsyth in press)

 

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